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Kira Yarmysh, Navalny's spokeswoman, wrote on Twitter that he had been hospitalised on Sunday morning with "severe swelling of the face and skin redness". She said the cause of Navalny's allergic reaction was unknown and that he had never suffered from such reactions in the past.

"It's still unclear what's wrong with him, but it looks strange," Navalny's lawyer, Olga Mikhailova, wrote on Facebook on Sunday evening. She said Navalny had had "an acute allergic reaction to an unknown allergen and swelling of the face" and insisted that he had had no allergies in his lifetime.

The Moscow hospital where Navalny's spokeswoman said he was being treated could not be reached for comment.

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In a separate incident on Sunday, Russian activist Dmitry Gudkov, who was among the opposition candidates barred from running in local elections this year, said he had been detained and taken to a Moscow police station.

The reason for Gudkov's detention was not immediately clear, his spokesman Alexei Obukhov told Reuters. Russia's Interior Ministry did not respond to a request for comment on Navalny and Gudkov's detention.

Obukhov said Gudkov, a former MP who challenged pro-Kremlin initiatives, had been detained as he walked out of a shop near his home, where he had been buying food for the protesters still being held. Navalny, Russia's most prominent opposition figure, has served several stints in jail in recent years for organising anti-government demonstrations.

The European Court of Human Rights last year ruled that Russia's arrests and detention of Navalny in 2012 and 2014 were politically motivated and breached his human rights, a rulin

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