Pygmy Sperm Whale
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^ Clarke, M.R. (2003). "Production and control of sound by the small sperm whale, Kogia breviceps and K. sima and their implications for other Cetacea". Journal of the Marine Biological Association of the United Kingdom . 83 (2): 241–263. doi : 10.1017/S0025315403007045h .
^ Jump up to: a b c d e f g h i R., Karol; C., Litchfield; D., Caldwell; M., Caldwell (1978). Compositional topography of melon and spermaceti organ lipids in the pygmy sperm whale Kogia breviceps: Implications for echolocation . Marine Biology , Volume 47 (2)
^ Cranford, T.W.; et al. (1996). "Functional morphology and homology in the odontocete nasal complex: implications for sound generation". Journal of Morphology . 228 (2): 223–285. doi : 10.1002/(SICI)1097-4687(199606)228:3<223::AID-JMOR1>3.0.CO;2-3 . PMID 8622183 .
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^ Poth, Fung, Gunturkun, Ridgway (2005-02-25). "Neuron numbers in sensory corticies of five delphinids compared to a physterid, the pygmy perm whale". Brain Research Bulletin . 66 (357–360). CS1 maint: multiple names: authors list ( link )
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^ Huckstadt, L.A. & Antezana, T. (2001). "An observation of parturition in a stranded Kogia breviceps ". Marine Mammal Science . 17 (2): 362–365. doi : 10.1111/j.1748-7692.2001.tb01277.x .
^ Willis, P.M. & Baird, R.W. (1998). "Status of the dwarf sperm whale, Kogia simus , with special reference to Canada" . Canadian Field-Naturalist . 112 (1): 114–125.
^ Thomas, J.A.; et al. (1990). "A new sound from a stranded pygmy sperm whale" (PDF) . Aquatic Mammals . 16 (1): 28–30.
^ Jump up to: a b Beatson, Emma (21 April 2007). "The diet of pygmy perm whales, Kogia breviceps, stranded in New Zealand: implications for conservation". Rev Fish Biol Fisheries . 10 (1007).
^ Long, D.J. (1991). "Apparent predation by a white shark Carcharadon charcharias on a pygmy sperm whale Kogia breviceps " (PDF) . Fishery Bulletin . 89 (3): 538–540.
^ Dunphy-Daly, M.M.; et al. (2008). "Temporal variation in dwarf sperm whale (Kogia sima) habitat use and group size off Great Abaco Island, Bahamas" . Marine Mammal Science . 24 (1): 171–182. doi : 10.1111/j.1748-7692.2007.00183.x .
^ "Pygmy sperm whale ( Kogia breviceps )" . NOAA Fisheries Office of Protected Resources web site . NOAA . 2014-10-20. Archived from the original on 2013-10-03 . Retrieved 2015-01-11 .
^ ХОРОШИЕ НОВОСТИ ПРО ЖИВОТНЫХ. 2017. НА КУНАШИРЕ (КУРИЛЫ) СПАСЛИ ЗАСТРЯВШЕГО НА МЕЛИ РЕДКОГО КАШАЛОТА С ДЕТЕНЫШЕМ . Retrieved on September 26, 2017
^ 2017. НА КУНАШИРЕ (КУРИЛЫ) СПАСЛИ ЗАСТРЯВШЕГО НА МЕЛИ РЕДКОГО КАШАЛОТА С ДЕТЕНЫШЕМ . Retrieved on September 26, 2017
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^ Davis, Lance E.; Gallman, Robert E.; Gleiter, Karin (January 1997). In Pursuit of Leviathan: Technology, Institutions, Productivity, and Profits in American Whaling, 1816-1906 . University of Chicago Press. pp. 20–56. ISBN 978-0-226-13789-6 .
^ Davis, Lance E.; Gallman, Robert E.; Gleiter, Karin (January 1997). In Pursuit of Leviathan: Technology, Institutions, Productivity, and Profits in American Whaling, 1816-1906 . University of Chicago Press. pp. 20–56. ISBN 978-0-226-13789-6 .
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^ Loranger, Linda (3 December 1991). "Pygmy Sperm Whale Dies In Mystic After Record Time In Captivity" . Hartford Courant . Retrieved 22 September 2016 .
^ Klingener, Nancy (2 November 2000). "Pygmy Whale Snags Fence, Dies After Four Months In Captivity" . Sun Sentinel . Retrieved 22 September 2016 .
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^ Official website of the Agreement on the Conservation of Small Cetaceans of the Baltic, North East Atlantic, Irish and North Seas
^ Official website of the Agreement on the Conservation of Cetaceans in the Black Sea, Mediterranean Sea and Contiguous Atlantic Area
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^ Official webpage of the Memorandum of Understanding for the Conservation of Cetaceans and Their Habitats in the Pacific Islands Region
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The pygmy sperm whale ( Kogia breviceps ) is one of two extant species in the family Kogiidae in the sperm whale superfamily . They are not often sighted at sea, and most of what is known about them comes from the examination of stranded specimens.
The pygmy sperm whale was first described by naturalist Henri Marie Ducrotay de Blainville in 1838. He based this on the head of an individual washed up on the coasts of Audierne in France in 1784, which was then stored in the Muséum d'histoire naturelle . He recognized it as a type of sperm whale and assigned it to the same genus as the sperm whale ( Physeter macrocephalus ) as Physeter breviceps . He noted its small size and nicknamed it " cachalot a tête courte "–small-headed sperm whale; further, the species name breviceps is Latin for "short-headed". [3] In 1846, zoologist John Edward Gray erected the genus Kogia for the pygmy sperm whale as Kogia breviceps , and said it was intermediate between the sperm whale and dolphins. [4]
In 1871, mammalogist Theodore Gill assigned it and Euphysetes (now the dwarf sperm whale , Kogia sima ) to the subfamily Kogiinae , and the sperm whale to the subfamily Physeterinae . [5] Both have now been elevated to the family level. In 1878, naturalist James Hector synonymized the dwarf sperm whale with the pygmy sperm whale, with both being referred to as K. breviceps until 1998. [6]
The pygmy sperm whale is not much larger than many dolphins . They are about 1.2 m (3 ft 11 in) at birth, growing to about 3.5 m (11 ft) at maturity. Adults weigh about 400 kg (880 lb). The underside is a creamy, occasionally pinkish colour and the back and sides are a bluish grey; however, considerable intermixing occurs between the two colours. The shark-like head is large in comparison to body size, given an almost swollen appearance when viewed from the side. A whitish marking, often described as a "false gill", is seen behind each eye. [7] [8]
The lower jaw is very small and slung low. The blowhole is displaced slightly to the left when viewed from above facing forward. The dorsal fin is very small and hooked; its size is considerably smaller than that of the dwarf sperm whale and may be used for diagnostic purposes.
Like its giant relative, the sperm whale , the pygmy sperm whale has a spermaceti organ in its forehead (see sperm whale for a discussion of its purpose). It also has a sac in its intestines that contains a dark red fluid. The whale may expel this fluid when frightened, perhaps to confuse and disorient predators. [9]
Dwarf and pygmy sperm whales possess the shortest rostrum of current day cetaceans with a skull that is greatly asymmetrical [10]
Pygmy sperm whales have from 50 to 55 vertebrae , and from 12 to 14 ribs on either side, although the latter are not necessarily symmetrical, and the hindmost ribs do not connect with the vertebral column. Each of the flippers has seven carpals , and a variable number of phalanges in the digits, reportedly ranging from two in the first digit to as many as 10 in the second digit. No true innominate bone exists; it is replaced by a sheet of dense connective tissue . The hyoid bone is unusually large, and presumably has a role in the whale's suction feeding. [8]
The pygmy sperm has between 20 and 32 teeth, all of which are set into the rostral part of the lower jaw. [11] Unusually, adults lack enamel due to a mutation in the enamelysin gene , [12] although enamel is present in very young individuals. [8]
Like other toothed whales , the pygmy sperm whale has a " melon ", a body of fat and wax in the head that it uses to focus and modulate the sounds it makes. [13] The inner core of the melon has a higher wax content than the outer cortex. The inner core transmits sound more slowly than the outer layer, allowing it to refract sound into a highly directional beam. [14] Behind the melon, separated by a thin membrane, is the spermaceti organ . Both the melon and the spermaceti organ are encased in a thick fibrous coat, resembling a bursa . [15] The whale produces sound by moving air through the right nasal cavity, which includes a valvular structure, or museau de singe , with a thickened vocal reed, functioning like the vocal cords of humans.
The stomach has three chambers. The first chamber, or forestomach, is not glandular, and opens directly into the second, fundic chamber, which is lined by digestive glands. A narrow tube runs from the second to the third, or pyloric, stomach, which is also glandular, and connects, via a sphincter , to the duodenum . Although fermentation of food material apparently occurs in the small intestine , no caecum is present. [16]
The rostroventral dura of the brain contains a significant concentration of magnetite crystals, which suggests that K. breviceps can navigate by magnetoreception . [8]
Studies have also shown that compared to the sperm whale , the pygmy sperm whale brain has significantly fewer neurons, which may be connected to a decreased complexity in social interaction and group-based living. [17]
Like all toothed whales, the pygmy sperm whale hunts prey by echolocation . Sound produced for echolocation by many odontocetes come in the form of high-frequency clicks. [18] The frequencies it uses are mostly ultrasonic, [14] peaking around 125 kHz. [18] The clicks from their echolocation has been recorded to last an average of 600 microseconds. When closing in on prey, the click repetition rate starts at 20 Hz and then begins to rise when nearing the target. [18]
The pulse sounds that pygmy sperm whales make for echolocation are generated primarily from the museau de singe or monkey's muzzle, which is an anatomical structure located within the whale's skull that produces sound when air passes through its lips. [14] The sound from the museau de singe is transferred to the attached cone of the spermaceti organ. Unique from other odontocetes, the spermaceti organ contacts the posterior of the whale's melon. [14] Fat from the core of the spermaceti organ helps direct sonic energy from the museau de singe to the melon. [14] The melon acts as a sonic magnifier and gives directionality to sound pulses, making echolocation more efficient. Fat on the interior of the melon has a lower molecular weight lipid than the surrounding outer melon. [14] Since the sound waves move from a lower velocity material to a higher one during sound production, the sound undergoes inward refraction and becomes increasingly focused. Variation in fat density within the melon ultimately contributes to the production of highly directional, ultrasonic sound beams in front of the melon. The combined melon and spermaceti organ system cooperate to focus echolocative sounds.
Like in most odontocetes, the known echoreception apparatus used by the pygmy sperm whale is linked to the fat-filled lower mandibles within the skull. [14] However, compositional topography of the pygmy sperm whale's skull indicates abnormally large fatty jowls surrounding the mandibles, suggesting a more intricate echoreception apparatus. [14] Additionally, an unusual cushion structure, of porous and spongy texture, found behind the museau de singe has been hypothesized of being a possible "pressure receptor". [14] The positioning of this cushion structure in close proximity to the largest cavities closest to the museau de singe may suggest that it is a sound absorber used for echoreception.
Although firm details concerning pygmy sperm whale reproduction are limited, they are believed to mate from April to September in the Southern Hemisphere and March to August in the Northern Hemisphere. [8] These whales become sexually mature at age 4-5, and like virtually all mammals, are iteroparous (reproducing many times during their lives). Once a female whale is impregnated, the average gestation period lasts 9–11 months, and unusually for cetaceans , the female gives birth to a single calf head-first. [19] Newborn calves are about 1.2 metres (3 ft 11 in) in length, weighing 50 kg, and are weaned around one year of age. [8] They are believed to live up to age 23.
The whale makes very inconspicuous movements. It rises to the surface slowly, with little splash or blow, and remains there motionless for some time. In Japan , the whale was historically known as the "floating whale" because of this. Its dive is equally lacking in grand flourish - it simply drops out of view. The species has a tendency to back away from rather than approach boats. Breaching has been observed, but is not common. [ citation needed ]
Pygmy sperm whales are normally either solitary or found in pairs, [20] but have been seen in groups up to six. [ citation needed ] Dives have been estimated to last an average of 11 minutes, although longer dives up to 45 minutes have been reported. [8] The ultrasonic clicks of pygmy sperm whales range from 60 to 200 kHz, peaking at 125 kHz, [18] and the animals also make much lower-frequency "cries" at 1 to 2 kHz. [21]
Analysis of stomach contents suggests that pygmy sperm whales feed primarily on cephalopods , most commonly including bioluminescent species found in midwater environments. Most of the cephalopod hunting is known to be pelagic, and fairly shallow, within the first 100 m of the surface. [22] The most common prey are reported to include glass squid , and lycoteuthid and ommastrephid squid, although the whales also consume other squid, and octopuses . They have also been reported to eat some deep-sea shrimps , but, compared with dwarf sperm whales, relatively few fish. [8]
Pygmy sperm whales and dwarf sperm whales are unique among cetaceans in using a form of "ink" to evade predation in a manner similar to squid. Both species have a sac in the lower portion of their intestinal tracts that contains up to 12 liters of dark reddish-brown fluid, which can be ejected to confuse or discourage potential predators. [25]
Pygmy sperm whales are found throughout the tropical and temperate waters of the Atlantic , Pacific , and Indian Oceans , [8] and occasionally among colder waters such as off Russia. [26] [27] However, they are rarely sighted at sea, so most data come from stranded animals - making a precise range and migration map difficult. They are believed to prefer off-shore waters, and are most frequently sighted in waters ranging from 400 to 1,000 m (1,300 to 3,300 ft) in depth, especially where upwelling water produces local concentrations of zooplankton and animal prey. [28] Their status is usually described as rare, but occasional patches of higher-density strandings suggest they might be more common than previously estimated. The total population is unknown.
Fossils
https://en.wikipedia.org/wiki/Pygmy_sperm_whale
https://savethewhales.org/pygmy-sperm-whale/
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Pygmy Sperm Whale
