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In , a book claiming that plants were sentient beings that feel emotions, prefer classical music to rock and roll, and can respond to the unspoken thoughts of humans hundreds of miles away landed on the New York Times best-seller list for nonfiction. The most memorable passages described the experiments of a former C. To his astonishment, Backster found that simply by imagining the dracaena being set on fire he could make it rouse the needle of the polygraph machine, registering a surge of electrical activity suggesting that the plant felt stress. Backster and his collaborators went on to hook up polygraph machines to dozens of plants, including lettuces, onions, oranges, and bananas. He claimed that plants reacted to the thoughts good or ill of humans in close proximity and, in the case of humans familiar to them, over a great distance. In one experiment designed to test plant memory, Backster found that a plant that had witnessed the murder by stomping of another plant could pick out the killer from a lineup of six suspects, registering a surge of electrical activity when the murderer was brought before it. Some had a stressful response when an egg was cracked in their presence, or when live shrimp were dropped into boiling water, an experiment that Backster wrote up for the International Journal of Parapsychology , in But the book had made its mark on the culture. Americans began talking to their plants and playing Mozart for them, and no doubt many still do. This might seem harmless enough; there will probably always be a strain of romanticism running through our thinking about plants. Luther Burbank and George Washington Carver both reputedly talked to, and listened to, the plants they did such brilliant work with. The authors pointed out that electrical and chemical signalling systems have been identified in plants which are homologous to those found in the nervous systems of animals. They also noted that neurotransmitters such as serotonin, dopamine, and glutamate have been found in plants, though their role remains unclear. It is only human arrogance, and the fact that the lives of plants unfold in what amounts to a much slower dimension of time, that keep us from appreciating their intelligence and consequent success. Plants dominate every terrestrial environment, composing ninety-nine per cent of the biomass on earth. Many plant scientists have pushed back hard against the nascent field, beginning with a tart, dismissive letter in response to the Brenner manifesto, signed by thirty-six prominent plant scientists Alpi et al. Santa Cruz and one of the signers of the Alpi letter, told me. The controversy is less about the remarkable discoveries of recent plant science than about how to interpret and name them: whether behaviors observed in plants which look very much like learning, memory, decision-making, and intelligence deserve to be called by those terms or whether those words should be reserved exclusively for creatures with brains. No one I spoke to in the loose, interdisciplinary group of scientists working on plant intelligence claims that plants have telekinetic powers or feel emotions. Nor does anyone believe that we will locate a walnut-shaped organ somewhere in plants which processes sensory data and directs plant behavior. Much of the research on plant intelligence has been inspired by the new science of networks, distributed computing, and swarm behavior, which has demonstrated some of the ways in which remarkably brainy behavior can emerge in the absence of actual brains. A slight, bearded Calabrian in his late forties, he comes across more like a humanities professor than like a scientist. When I visited him earlier this year at the International Laboratory of Plant Neurobiology, at the University of Florence, he told me that his conviction that humans grossly underestimate plants has its origins in a science-fiction story he remembers reading as a teen-ager. The aliens proceed ruthlessly to exploit us. It creates a resilience. Indeed, many of the most impressive capabilities of plants can be traced to their unique existential predicament as beings rooted to the ground and therefore unable to pick up and move when they need something or when conditions turn unfavorable. A highly developed sensory apparatus is required to locate food and identify threats. The sensory capabilities of plant roots fascinated Charles Darwin, who in his later years became increasingly passionate about plants; he and his son Francis performed scores of ingenious experiments on plants. Scientists have since found that the tips of plant roots, in addition to sensing gravity, moisture, light, pressure, and hardness, can also sense volume, nitrogen, phosphorus, salt, various toxins, microbes, and chemical signals from neighboring plants. Roots about to encounter an impenetrable obstacle or a toxic substance change course before they make contact with it. Roots can tell whether nearby roots are self or other and, if other, kin or stranger. Normally, plants compete for root space with strangers, but, when researchers put four closely related Great Lakes sea-rocket plants Cakile edentula in the same pot, the plants restrained their usual competitive behaviors and shared resources. Davis, explained, when I asked him for an example of plant decision-making. Many drugs, from aspirin to opiates, derive from compounds designed by plants. Unable to run away, plants deploy a complex molecular vocabulary to signal distress, deter or poison enemies, and recruit animals to perform various services for them. A recent study in Science found that the caffeine produced by many plants may function not only as a defense chemical, as had previously been thought, but in some cases as a psychoactive drug in their nectar. The caffeine encourages bees to remember a particular plant and return to it, making them more faithful and effective pollinators. One of the most productive areas of plant research in recent years has been plant signalling. Sometimes this warning signal contains information about the identity of the insect, gleaned from the taste of its saliva. When antelopes browse acacia trees, the leaves produce tannins that make them unappetizing and difficult to digest. When food is scarce and acacias are overbrowsed, it has been reported, the trees produce sufficient amounts of toxin to kill the animals. Perhaps the cleverest instance of plant signalling involves two insect species, the first in the role of pest and the second as its exterminator. Several species, including corn and lima beans, emit a chemical distress call when attacked by caterpillars. Parasitic wasps some distance away lock in on that scent, follow it to the afflicted plant, and proceed to slowly destroy the caterpillars. The first important discoveries in plant communication were made in the lab in the nineteen-eighties, by isolating plants and their chemical emissions in Plexiglas chambers, but Rick Karban, the U. Davis ecologist, and others have set themselves the messier task of studying how plants exchange chemical signals outdoors, in a natural setting. On a sun-flooded hillside high in the Sierras, he introduced me to the ninety-nine sagebrush plants—low, slow-growing gray-green shrubs marked with plastic flags—that he and his colleagues have kept under close surveillance for more than a decade. Karban, a fifty-nine-year-old former New Yorker, is slender, with a thatch of white curls barely contained by a floppy hat. He has shown that when sagebrush leaves are clipped in the spring—simulating an insect attack that triggers the release of volatile chemicals—both the clipped plant and its unclipped neighbors suffer significantly less insect damage over the season. Karban believes that the plant is alerting all its leaves to the presence of a pest, but its neighbors pick up the signal, too, and gird themselves against attack. He found that the more closely related the plants the more likely they are to respond to the chemical signal, suggesting that plants may display a form of kin recognition. Helping out your relatives is a good way to improve the odds that your genes will survive. The field work and data collection that go into making these discoveries are painstaking in the extreme. At the bottom of a meadow raked by the slanted light of late summer, two collaborators from Japan, Kaori Shiojiri and Satomi Ishizaki, worked in the shade of a small pine, squatting over branches of sagebrush that Karban had tagged and cut. Using clickers, they counted every trident-shaped leaf on every branch, and then counted and recorded every instance of leaf damage, one column for insect bites, another for disease. At the top of the meadow, another collaborator, James Blande, a chemical ecologist from England, tied plastic bags around sagebrush stems and inflated the bags with filtered air. After waiting twenty minutes for the leaves to emit their volatiles, he pumped the air through a metal cylinder containing an absorbent material that collected the chemical emissions. At the lab, a gas chromatograph-mass spectrometer would yield a list of the compounds collected—more than a hundred in all. Blande offered to let me put my nose in one of the bags; the air was powerfully aromatic, with a scent closer to aftershave than to perfume. Research on plant communication may someday benefit farmers and their crops. Plant-distress chemicals could be used to prime plant defenses, reducing the need for pesticides. Karban told me that, in the nineteen-eighties, people working on plant communication faced some of the same outrage that scientists working on plant intelligence a term he cautiously accepts do today. People would literally be screaming at one another at scientific meetings. The meeting consisted of three days of PowerPoint presentations delivered in a large, modern lecture hall at the University of British Columbia before a hundred or so scientists. Most of the papers were highly technical presentations on plant signalling—the kind of incremental science that takes place comfortably within the confines of an established scientific paradigm, which plant signalling has become. But a handful of speakers presented work very much within the new paradigm of plant intelligence, and they elicited strong reactions. Gagliano, who is tall, with long brown hair parted in the middle, based her experiment on a set of protocols commonly used to test learning in animals. When the fernlike leaves of the mimosa are touched, they instantly fold up, presumably to frighten insects. The mimosa also collapses its leaves when the plant is dropped or jostled. Gagliano potted fifty-six mimosa plants and rigged a system to drop them from a height of fifteen centimetres every five seconds. She reported that some of the mimosas started to reopen their leaves after just four, five, or six drops, as if they had concluded that the stimulus could be safely ignored. Was it just fatigue? Apparently not: when the plants were shaken, they again closed up. Even after twenty-eight days, the lesson had not been forgotten. She reminded her colleagues that, in similar experiments with bees, the insects forgot what they had learned after just forty-eight hours. A lively exchange followed. Gagliano pointed out that electric shock, an equally artificial trigger, is often used in animal-learning experiments. Another scientist suggested that perhaps her plants were not habituated, just tuckered out. She argued that twenty-eight days would be plenty of time to rebuild their energy reserves. On my way out of the lecture hall, I bumped into Fred Sack, a prominent botanist at the University of British Columbia. At lunch, I sat with a Russian scientist, who was equally dismissive. Later that afternoon, Gagliano seemed both stung by some of the reactions to her presentation and defiant. Adaptation is far too slow a process to explain the behavior she had observed, she told me. Rick Karban consoled Gagliano after her talk. The system is just not ready. Scientists are often uncomfortable talking about the role of metaphor and imagination in their work, yet scientific progress often depends on both. Yet the introduction of the term has raised a series of questions and inspired a set of experiments that promise to deepen our understanding not only of plants but potentially also of brains. Mancuso is the poet-philosopher of the movement, determined to win for plants the recognition they deserve and, perhaps, bring humans down a peg in the process. His somewhat grandly named International Laboratory of Plant Neurobiology, a few miles outside Florence, occupies a modest suite of labs and offices in a low-slung modern building. Here a handful of collaborators and graduate students work on the experiments Mancuso devises to test the intelligence of plants. Giving a tour of the labs, he showed me maize plants, grown under lights, that were being taught to ignore shadows; a poplar sapling hooked up to a galvanometer to measure its response to air pollution; and a chamber in which a PTR-TOF machine—an advanced kind of mass spectrometer—continuously read all the volatiles emitted by a succession of plants, from poplars and tobacco plants to peppers and olive trees. He is also gentle and unassuming, even when what he is saying is outrageous. One of his experiments was carried on board the last flight of the space shuttle Endeavor, in May of A decade ago, Mancuso persuaded a Florentine bank foundation to underwrite much of his research and help launch the Society for Plant Neurobiology; his lab also receives grants from the European Union. It turns out that I am not alone: philosophers and psychologists have been arguing over the definition of intelligence for at least a century, and whatever consensus there may once have been has been rapidly slipping away. Most definitions of intelligence fall into one of two categories. The first is worded so that intelligence requires a brain; the definition refers to intrinsic mental qualities such as reason, judgment, and abstract thought. Not surprisingly, the plant neurobiologists jump into this second camp. The idea remains unproved and controversial. The hypothesis that intelligent behavior in plants may be an emergent property of cells exchanging signals in a network might sound far-fetched, yet the way that intelligence emerges from a network of neurons may not be very different. That sense we get when we think about what might govern a plant—that there is no there there, no wizard behind the curtain pulling the levers—may apply equally well to our brains. Next came Darwin, who brought the humbling news that we are the product of the same natural laws that created animals. In the last century, the formerly sharp lines separating humans from animals—our monopolies on language, reason, toolmaking, culture, even self-consciousness—have been blurred, one after another, as science has granted these capabilities to other animals. Plant neurobiologists have chosen to define intelligence democratically, as an ability to solve problems or, more precisely, to respond adaptively to circumstances, including ones unforeseen in the genome. We exist on a continuum with the acacia, the radish, and the bacterium. I asked him why he thinks people have an easier time granting intelligence to computers than to plants. We tend to think of memories as immaterial, but in animal brains some forms of memory involve the laying down of new connections in a network of neurons. In plants, it has long been known that experiences such as stress can alter the molecular wrapping around the chromosomes; this, in turn, determines which genes will be silenced and which expressed. More recently, scientists have found that life events such as trauma or starvation produce epigenetic changes in animal brains coding for high levels of cortisol, for example that are long-lasting and can also be passed down to offspring, a form of memory much like that observed in plants. At one point, he told me about the dodder vine, Cuscuta europaea , a parasitic white vine that winds itself around the stalk of another plant and sucks nourishment from it. Having selected a target, the vine then performs a kind of cost-benefit calculation before deciding exactly how many coils it should invest—the more nutrients in the victim, the more coils it deploys. I asked Mancuso whether he was being literal or metaphorical in attributing intention to plants. Time-lapse photography is perhaps the best tool we have to bridge the chasm between the time scale at which plants live and our own. This example was of a young bean plant, shot in the lab over two days, one frame every ten minutes. A metal pole on a dolly stands a couple of feet away. Each spring, I witness the same process in my garden, in real time. I always assumed that the bean plants simply grow this way or that, until they eventually bump into something suitable to climb. Mancuso speculates that the plant could be employing a form of echolocation. And it is striving there is no other word for it to get there: reaching, stretching, throwing itself over and over like a fly rod, extending itself a few more inches with every cast, as it attempts to wrap its curling tip around the pole. As soon as contact is made, the plant appears to relax; its clenched leaves begin to flutter mildly. All this may be nothing more than an illusion of time-lapse photography. Taiz also pointed out that the bean in the video was leaning toward the pole in the first frame. Mancuso then sent me another video with two perfectly upright bean plants that exhibited very similar behavior. Taiz was now intrigued. But this is one of the features of consciousness: You know your position in the world. A stone does not. But apparently not. Pain is adaptive. Unprepared to consider the ethical implications of plant intelligence, I could feel my resistance to the whole idea stiffen. Descartes, who believed that only humans possessed self-consciousness, was unable to credit the idea that other animals could suffer from pain. So he dismissed their screams and howls as mere reflexes, as meaningless physiological noise. Could it be remotely possible that we are now making the same mistake with plants? That the perfume of jasmine or basil, or the scent of freshly mowed grass, so sweet to us, is as the ecologist Jack Schultz likes to say the chemical equivalent of a scream? Lincoln Taiz has little patience for the notion of plant pain, questioning what, in the absence of a brain, would be doing the feeling. We can never determine with certainty whether plants feel pain or whether their perception of injury is sufficiently like that of animals to be called by the same word. Mancuso believes that, because plants are sensitive and intelligent beings, we are obliged to treat them with some degree of respect. That means protecting their habitats from destruction and avoiding practices such as genetic manipulation, growing plants in monocultures, and training them in bonsai. But it does not prevent us from eating them. He cited their modular structure and lack of irreplaceable organs in support of this view. The central issue dividing the plant neurobiologists from their critics would appear to be this: Do capabilities such as intelligence, pain perception, learning, and memory require the existence of a brain, as the critics contend, or can they be detached from their neurobiological moorings? The question is as much philosophical as it is scientific, since the answer depends on how these terms get defined. The proponents of plant intelligence argue that the traditional definitions of these terms are anthropocentric—a clever reply to the charges of anthropomorphism frequently thrown at them. Their attempt to broaden these definitions is made easier by the fact that the meanings of so many of these terms are up for grabs. Indeed, I found more consensus on the underlying science than I expected. To define certain words in such a way as to bring plants and animals beneath the same semantic umbrella—whether of intelligence or intention or learning—is a philosophical choice with important consequences for how we see ourselves in nature. Plant neurobiologists are intent on taking away our skyhook, completing the revolution that Darwin started but which remains—psychologically, at least—incomplete. Upon a foundation of the simplest competences—such as the on-off switch in a computer, or the electrical and chemical signalling of a cell—can be built higher and higher competences until you wind up with something that looks very much like intelligence. All species face the same existential challenges—obtaining food, defending themselves, reproducing—but under wildly varying circumstances, and so they have evolved wildly different tools in order to survive. Impressive as it is to us, self-consciousness is just another tool for living, good for some jobs, unhelpful for others. In addition to being a plant physiologist, Lincoln Taiz writes about the history of science. It implies free will. Of course, one could argue that humans lack free will too, but that is a separate issue. I asked Mancuso if he thought that a plant decides in the same way we might choose at a deli between a Reuben or lox and bagels. One way to exalt plants is by demonstrating their animal-like capabilities. But another way is to focus on all the things plants can do that we cannot. Without locomotion? By focussing on the otherness of plants rather than on their likeness, Mancuso suggested, we stand to learn valuable things and develop important new technologies. Mancuso was about to begin a collaboration with a prominent computer scientist to design a plant-based computer, modelled on the distributed computing performed by thousands of roots processing a vast number of environmental variables. His collaborator, Andrew Adamatzky, the director of the International Center of Unconventional Computing, at the University of the West of England, has worked extensively with slime molds, harnessing their maze-navigating and computational abilities. In an e-mail, Adamatzky said that, as a substrate for biological computing, plants offered both advantages and disadvantages over slime molds. If you want something swimming, you look at a fish. But what about imitating plants instead? What would that allow you to do? Explore the soil! Citing the research of Suzanne Simard, a forest ecologist at the University of British Columbia, and her colleagues, Mancuso showed a slide depicting how trees in a forest organize themselves into far-flung networks, using the underground web of mycorrhizal fungi which connects their roots to exchange information and even goods. When I reached Simard by phone, she described how she and her colleagues track the flow of nutrients and chemical signals through this invisible underground network. They injected fir trees with radioactive carbon isotopes, then followed the spread of the isotopes through the forest community using a variety of sensing methods, including a Geiger counter. Within a few days, stores of radioactive carbon had been routed from tree to tree. Every tree in a plot thirty metres square was connected to the network; the oldest trees functioned as hubs, some with as many as forty-seven connections. The diagram of the forest network resembled an airline route map. The evergreen species will tide over the deciduous one when it has sugars to spare, and then call in the debt later in the season. In his talk, Mancuso juxtaposed a slide of the nodes and links in one of these subterranean forest networks with a diagram of the Internet, and suggested that in some respects the former was superior. As I listened to Mancuso limn the marvels unfolding beneath our feet, it occurred to me that plants do have a secret life, and it is even stranger and more wonderful than the one described by Tompkins and Bird. When most of us think of plants, to the extent that we think about plants at all, we think of them as old—holdovers from a simpler, prehuman evolutionary past. But for Mancuso plants hold the key to a future that will be organized around systems and technologies that are networked, decentralized, modular, reiterated, redundant—and green, able to nourish themselves on light. So it will have a solid structure and a center, but that is inherently fragile. Save this story Save this story. Copy link to cartoon Copy link to cartoon. Link copied. Michael Pollan teaches journalism at the University of California, Berkeley. Annals of Medicine. The Trip Treatment. By Michael Pollan. At first, scientists just wanted to figure out the best way to kill these pests. Then they decided that studying rat society could reveal the future of our own. By Elizabeth Kolbert. Annals of Zoology. How Scientists Started to Decode Birdsong. Language is said to make us human. What if birds talk, too? By Rivka Galchen. Book Currents. The author reads his story from the October 21, , issue of the magazine. The Lede. Tim Walz and J. Duelling visions of fatherhood will define the Vice-Presidential debate. By Molly Fischer. Annals of Psychology. By Eren Orbey. The New Yorker Interview. In her new Netflix special, the comedian turns a tragic life episode into a riotous study of motherhood, mortality, and the meaning of pet heaven. By Alexandra Schwartz. The New Yorker Documentary. The documentary short by Patrick Bresnan and Ivete Lucas follows Michael Mullen on his rounds attending to pets and their owners. Briefly Noted. The Financial Page. A new book by two New York Times investigative reporters comprehensively debunks the notion that Trump is a good businessman. By John Cassidy. Pop Music. Sophie Is Gone. Her Music Lives On. By Jia Tolentino.

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